te = the; pradhāneṣu = of the chief; yūtheṣu = groups; harīṇām = of monkeys; hari-yūthapāḥ = leaders of monkeys [from Sugrīva to Suṣeṇa]; babhūvuḥ = became; yūthapa-śreṣṭhāḥ = chiefs of the leaders; vīrān = to heroic; ajanayan = they gave birth; harīn ca = monkeys.